seemab akram, shamsul khamis, shahrizim zulkifly, rishzuan talib, nurul izza ab ghani.  (2023).  phylogeny study of 20 selected species of zingiberaceae from ex situ collections in peninsular malaysia.  - pertanika j. trop. agric. sci..  329-345. 

muhammad izzelen izzauddin mamat, shamsul khamis, muhammad faqih luqman fuaad, nor zalipah mohamed, chik maslinda omar, daria mathew abdullah, wahizatul afzan azm.  (2023).  pollen sources foraged by domesticated stingless bee (heterotrigona itama) reared in gelam forests of terengganu, malaysia.  - serangga.  120-138. 

n.h.a. kadir, w.m.n.h.w. salleh, n.a. ghani, s. khamis, m.a.a juhairi.  (2023).  chemical composition and acetylcholinesterase activity of syzygium pyrifolium (blume) dc. essential oil.  - latin american applied research.  357-359. 

ahmad azrul hakim yahaya, wan mohd nuzul hakimi wan salleh, nurunajah ab ghani, shamsul khamis, nurul syafiqah rezali, mohd afiq aizat juhari.  (2022).  chemical composition and bioactivities of magnolia candollii h.keng essential oil.  - zeitschrift fur naturforschung.  519-523. 

muhamad faris osman, siti munirah mohd faudzi, shamsul khamis, shahrul razid sarbini, khozirah shaari.  (2022).  shedding light on pipers identity via computational mass spectrometry.  - malaysian journal of chemistry.  150-160. 

nur-fatehah sr, nur-shafikah i, shamsul k, wan-juliana wa, norela sulaiman, shahril mh, nurul-darsani ad.  (2022).  stand structure and productivity of the firefly riverine mangrove habitat at three rivers in peninsular malaysia.  - journal of tropical forest science.  114-126. 

siti nurfaeiza abd razak, wan juliana wan ahmad, shamsul khamis, shukor md nor.  (2022).  morpho-physiological strategies of shorea leprosula miq. and shorea acuminata dyer in response to light intensity and nutrient amendments.  - forest.  1-18. 

jumaat h. adam, mohd afiq aizat-juhari, nik nadira-farhana nik ariff, shamsul khamis, mohd hizamri mohd yasin, kiew-lian wan.  (2022).  rafflesia tunku-azizahiae (rafflesiaceae), a new species from pahang, malaysia.  - sains malaysiana.  3843-3855. 

a. r. siti nurfaeiza, k. shamsul, w.a. wan juliana, m.n. shukor.  (2022).  tropical rainforest regeneration in an area devastated by dam impoundment in hulu terengganu, malaysia.  - the malaysian forester.  96-108. 

syarifah haniera sheikh kamal, mohd nazip suratman, shamsul khamis, ahmad najmi nik hassan, mohd syaiful mohammad.  (2022).  growth rate, mortality rate and life cycle of rafflesia azlanii and r. cantleyi in belum-temenggor forest complex, perak, malaysia.  - sains malaysiana.  943-957. 

anisa s. al-hakimi, sarah abdul razak, n. omar, abdulwahab a. saeed, shamsul khamis, a. latiff.  (2022).  stem and leaf anatomical studies of selected species of barlerieae and ruellieae (acanthaceae) from yemen.  - sains malaysiana.  1609-1624. 

wan mohd nuzul hakimi wan salleh, natasa mohd shakri, shamsul khamis, william n. setzer, muhammad helmi nadri.  (2022).  chemical composition of three malaysian horsfieldia essential oils.  - natural product research.  1909-1913. 

m.a.m. azhar, w.m.n.h.w. salleh, s. khamis, n.a. ghani.  (2022).  variation in essential oil composition of three litsea species from malaysia.  - rivista italiana delle sostanze grasse.  57-61. 

w.m.n.h.w. salleh, n.m. shakri, m.a. nafiah, s. khamis.  (2022).  chemical constituents and acetylcholinesterase activity from polyalthia sumatrana (miq.) kurz.  - latin american applied research.  73-76. 

rasyidah wahab, ahmad asnawi mus, kartini saibeh, shamsul khamis, handry mujih, geofarry gunsalam, dasini, ezron gerald, juhinin sariman, qarin aaron muhammad, qaid ahmad muhammad.  (2021).  soil physico-chemistry in the habitat of rafflesia in kinabalu park, sabah, malaysia..  - journal of tropical biology and conservation.  149-165. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, hakimi kassim, alene tawang.  (2021).  chemical composition of the essential oil of dysoxylum cauliflorum hiern (meliaceae).  - natural volatiles & essential oils.  7-12. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, hakimi kassim, alene tawang.  (2021).  essential oil composition of strychnos axillaris colebr. (loganiaceae).  - natural volatiles & essential oils.  13-17. 

a.k.n. hayati, k. shamsul, w.a.w. juliana, f.s. mohd. taib, m.n. shukor, m.h. shahril, e.a. filza.  (2021).  habitat and edaphic preference of rafflesia r. br. plant communities in royal belum state park, perak, peninsular malaysia.  - journal of environmental biology.  865-871. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2021).  composition of essential oil from actinodaphne sesquipedalis and its lipoxygenase activity.  - chemistry of natural compounds.  553-555. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2021).  chemical composition of the essential oil of diospyros argentea.  - chemistry of natural compounds.  556-557. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2021).  composition of volatile oil from anaxagorea javanica.  - chemistry of natural compounds.  558-559. 

syarifah haniera sheikh kamal, mohd nazip suratman, shamsul khamis, ahmad najmi nik hassan, mohd syaiful mohammad.  (2021).  host-parasitic relationships between tetrastigma rafflesiae and rafflesia azlanii and rafflesia cantleyi in belum-temenggor forest complex, perak, malaysia.  - pertanika journal of tropical agricultural science.  755-771. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, muhammad nurakmal abdul rahman, mohd azlan nafiah.  (2021).  chemical composition and acetylcholinesterase activity of the essential oil of anisophyllea disticha.  - chemistry of natural compounds.  371-373. 

muhamad faris osman, soo yee lee, shahrul razid sarbini, siti munirah mohd faudzi, shamsul khamis, badrul hisyam zainudin, khozirah shaari.  (2021).  metabolomics-driven discovery of an introduced species and two malaysian piper betle l. variants.  - plants.  1-19. 

muhammad ammar mohd azhar, wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2021).  essential oil composition of beilschmiedia insignis from malaysia.  - chemistry of natural compounds.  374-375. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, salam ahmed abed.  (2021).  characterization of volatile components of chassalia chartacea and its acetylcholinesterase inhibitory activity.  - chemistry of natural compounds.  376-377. 

nor shafikah idris, muzzneena ahmad mustapha, norela sulaiman, shamsul khamis, shahril mohd husin, nurul darsani ahmad darbis.  (2021).  the dynamics of landscape changes surrounding a firefly ecotourism area.  - global ecology and conservation.  1-20. 

wan mohd nuzul hakimi wan salleh, mohd azlan nafiah, shamsul khamis, salam ahmed abed.  (2021).  chemical composition and lipoxygenase inhibitory activity of alseodaphne peduncularis meisn. essential oil.  - nat. volatiles & essent. oils.  29-33. 

siti nur fatehah, r., shamsul, k., wan juliana, w. a., norela sulaiman, md. shahril,m. h., a. d. nurul darsani.  (2021).  distribution patterns of plant communities in riverine mangrove habitats and their associations with conductivity factors in the west coast of peninsular malaysia.  - the malaysian forester.  169-184. 

natasa mohd shakri, wan mohd nuzul hakimi wan salleh, shamsul khamis, nor azah mohamad ali, muhammad helmi nadri.  (2020).  composition of the essential oils of three malaysian xylopia species (annonaceae).  - zeitschrift fur naturforschung c a journal of biosciences.  479-484. 

natasa mohd shakri, wan mohd nuzul hakimi wan salleh, shamsul khamis, nor azah mohamad ali.  (2020).  chemical characterization of goniothalamus macrophyllus and goniothalamus malayanus leaves essential oils.  - zeitschrift fur naturforschung c a journal of biosciences.  485-488. 

natasa mohd shakri, wan mohd nuzul hakimi wan salleh, shamsul khamis, nor azah mohamad ali, shazlyn milleana shaharudin.  (2020).  chemical composition of the essential oils of four polyalthia species from malaysia.  - zeitschrift fuer naturforschung, section c: a journal of biosciences.  473-478. 

shamsul, k., nik norafida, n. a., mat salleh, k. and a. latiff.  (2020).  tree species composition, diversity and biomass estimation in lower montane forest at fraser's hill, pahang, peninsular malaysia.  - the malaysian forester.  114-127. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2020).  chemical composition of the essential oil of diospyros wallichii king & gamble (ebenaceae).  - natural volatiles & essential oils.  12-17. 

muhammad ammar mohd azhar, wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2020).  essential oil composition of three cryptocarya species from malaysia.  - zeitschrift fur naturforschung section c-a journal of biosciences.  297-301. 

a.k. nur hayati, k. shamsul, w.a. wan juliana, m. n. shukor, m. h. shahril, e. alyaa filza.  (2020).  plant community structure and diversity of the rafflesia habitat at the royal belum state park, perak, malaysia.  - the malaysian forester.  387-404. 

sani aliyu, norazah basar1, yakubu rufai, shamsul khamis.  (2020).  phytochemical analysis and isolation of secondary metabolites from persea declinata.  - malaysian journal of analytical sciences.  970-977. 

wan mohd nuzul hakimi wan salleh, nur kamilah mohammed, nur athirah hashim, shamsul khamis, salam ahmed abed, saidu jibril.  (2020).  chemical constituents of beilschmiedia penangiana.  - chemistry of natural compounds.  576-577. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2020).  chemical composition and lipoxygenase activity of the leaves essential oil of rothmannia macrophylla (hook.f.) bremek.  - journal of essential oil bearing plants.  331-336. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2020).  chemical composition and anticholinesterase inhibitory activity of pavetta graciliflora wall. ex ridl. essential.  - zeitschrift fur naturforschung c a journal of biosciences.  467-471. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, muhammad nurakmal abdul rahman, mohd azlan nafiah.  (2020).  chemical composition and biological activities of dipterocarpus cornutus dyer essential oil.  - journal of biosciences: zeitschrift für naturforschung c.  171-175. 

wan mohd nuzul hakimi wan salleh, norkamilah abdullah, nur athirah hashim, heng yen khong, shamsul khamis.  (2019).  aporphine alkaloids from piper erecticaule and acetylcholinesterase inhibitory activity.  - boletin latinoamericano y del caribe de plantas medicinales y aromaticas.  527-532. 

wan mohd nuzul hakimi wan salleh, nur athirah hashim, shamsul khamis.  (2019).  chemical constituents and lipoxygenase inhibitory activity of piper stylosum miq.  - bulletin of the chemical society of ethiopia.  587-592. 

shamsul k., m. s. nizam, anisa s. al-hakimi, n. talip.  (2019).  taxonomic value of petiole anatomy in malaysian litsea complex (lauraceae).  - malayan nature journal.  453-465. 

nur athirah hashim, farediah ahmad, wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2019).  a new amide from piper maingayi hk.f. (piperaceae).  - natural product communications.  1-5. 

muhamad ikhwanuddin mat esa, rosimah nulit, shamsul khamis, rusea go.  (2019).  ethnomedical survey of plants used by the bateq in kuala koh, kelantan, malaysia.  - malayan nature journal.  439-451. 

faridah qz, ayuni m. n, shamsul khamis, anisa s. al-hakimi, rosimah nulit, norhidayah mad halid.  (2019).  traditional knowledge on aromatic plants as utilised by the jahai tribe in royal belum state park, perak, malaysia.  - malayan nature journal.  1-8. 

nur hayati a. k, shamsul k., wan juliana w. a., shahril m. h., alyaa filza e., mohamed shah redza h..  (2019).  re-assessment of the rafflesia populations and habitats in royal belum state park, perak, peninsular malaysia.  - malayan nature journal.  493-498. 

nur-athirah abdullah, siti nur fatehah radzi, lailatul-nadhirah asri, nor shafikah idris, shahril husin, azman sulaiman, shamsul khamis, norela sulaiman, izfa riza hazmi.  (2019).  insect community in riparian zones of sungai sepetang, sungai rembau and sungai chukai of peninsular malaysia.  - biodiversity data journal.  1-24. 

faridah qamaruz zaman, anisa s. al-hakimi, shamsul khamis, fatin f. ruhaizin and syuhada. m. zaidi.  (2018).  anatomy and histochemistry of structures producing aroma in leaves of sygyzium aromaticum (l.) merr. and clausena excavata burm. f..  - pertanika journal of tropical agricultural science.  .