shah badri mohd nor, ahmad fariz mohamed, shamsul khamis.  (2025).  forest certification management experience in asia pacific and europe: a case study of malaysia and sweden.  - journal of forest science.  269 - 285. 

shamsul khamis, siti nurfaeiza abd razak, ras elmeera mohd rashid, loh khai vee, nadzifah yaakub, hafizan juahir, mohd nordin abdul rahman, jeyanny vijayanathan.  (2025).  tree species composition and community structure of vegetation on bris soil of coastal terengganu, peninsular malaysia.  - the malayan nature journal.  125 - 134. 

roberta chaya tawie tingga , millawati gani , nur azimah osman , nor rahman aifat , eddie chan , shamsul khamis , emelda rosseleena rohani , norlinda mohd-daut , abd rahman mohd-ridwan, badrul munir md-zain.  (2024).  interspecific variation in the diet of symphalangus syndactylus and macaca nemestrina at genting highlands, pahang, peninsular malaysia.  - biodiversity data journal.  1 - 18. 

haja maideen, norlaila najwa rahman, nur aliah, nurul nadhirah, noraini talip, nik norhazrina, nur aqilah bakray, ahmad fitri zohari, nur farah ain zainee, mohd nizam mohd said, shamsul khamis, nur syazwani mohamad basir.  (2024).  anatomical characteristics of selected dennstaedtiaceae species from peninsular malaysia.  - malaysian applied biology.  229 - 237. 

roberta chaya tawie tingga, millawati gani, nur azimah osman, nor rahman aifat, eddie chan, shamsul khamis, emelda rosseleena rohani, norlinda mohd-daut, abd rahman mohd ridwan, badrul munir md-zain.  (2024).  interspecific variation in the diet of symphalangus syndactylus and macaca nemestrina at genting highlands, pahang, peninsular malaysia.  - biodiversity data journal.  1-18. 

noraini talip, jayasrie eyamalay, mohamad ruzi abdul rahman, mohd nizam mohd said, haja maideen kader maideen, nik norhazrina nik mohd kamil, nur aqilah mustafa bakray, shamsul khamis, ahmad fitri zohari, nur farah ain zainee, hamidun bunawan.  (2024).  ciri adaptasi anatomi daun bougainvillea spectabilis w. yang dijangkiti virus bougainvillea chlorotic vein-banding.  - malaysian applied biology.  43-54. 

wan mohd nuzul hakimi wan salleh, mohd hafiz arzmi, shamsul khamis, mohd afiq aizat juhari.  (2024).  chemical composition and antioxidant activity of the essential oil of tabernaemontana dichotoma.  - chemistry of natural compounds.  169-170. 

nurunajah ab ghani, wan mohd nuzul hakimi wan salleh, abubakar siddiq salihu, nurulfazlina edayah rasol, shamsul khamis.  (2024).  essential oil composition of antidesma cuspidatum and acetylcholinesterase inhibitory activity.  - chemistry of natural compounds.  167-168. 

seemab akram, shamsul khamis, shahrizim zulkifly, rishzuan talib, nurul izza ab ghani.  (2023).  phylogeny study of 20 selected species of zingiberaceae from ex situ collections in peninsular malaysia.  - pertanika j. trop. agric. sci..  329-345. 

seemab akram, shahrizim zulkifly, shamsul khamis, nurul izza ab ghani.  (2023).  preliminary in silico analysis of chs1 gene in commelinids clade: family zingiberaceae, costaceae, and poaceae.  - pertanika j. trop. agric. sci..  425-437. 

n.h.a. kadir, w.m.n.h.w. salleh, n.a. ghani, s. khamis, m.a.a juhairi.  (2023).  chemical composition and acetylcholinesterase activity of syzygium pyrifolium (blume) dc. essential oil.  - latin american applied research.  357-359. 

muhammad izzelen izzauddin mamat, shamsul khamis, muhammad faqih luqman fuaad, nor zalipah mohamed, chik maslinda omar, daria mathew abdullah, wahizatul afzan azm.  (2023).  pollen sources foraged by domesticated stingless bee (heterotrigona itama) reared in gelam forests of terengganu, malaysia.  - serangga.  120-138. 

a. r. siti nurfaeiza, k. shamsul, w.a. wan juliana, m.n. shukor.  (2022).  tropical rainforest regeneration in an area devastated by dam impoundment in hulu terengganu, malaysia.  - the malaysian forester.  96-108. 

nur-fatehah sr, nur-shafikah i, shamsul k, wan-juliana wa, norela sulaiman, shahril mh, nurul-darsani ad.  (2022).  stand structure and productivity of the firefly riverine mangrove habitat at three rivers in peninsular malaysia.  - journal of tropical forest science.  114-126. 

syarifah haniera sheikh kamal, mohd nazip suratman, shamsul khamis, ahmad najmi nik hassan, mohd syaiful mohammad.  (2022).  growth rate, mortality rate and life cycle of rafflesia azlanii and r. cantleyi in belum-temenggor forest complex, perak, malaysia.  - sains malaysiana.  943-957. 

w.m.n.h.w. salleh, n.m. shakri, m.a. nafiah, s. khamis.  (2022).  chemical constituents and acetylcholinesterase activity from polyalthia sumatrana (miq.) kurz.  - latin american applied research.  73-76. 

m.a.m. azhar, w.m.n.h.w. salleh, s. khamis, n.a. ghani.  (2022).  variation in essential oil composition of three litsea species from malaysia.  - rivista italiana delle sostanze grasse.  57-61. 

wan mohd nuzul hakimi wan salleh, natasa mohd shakri, shamsul khamis, william n. setzer, muhammad helmi nadri.  (2022).  chemical composition of three malaysian horsfieldia essential oils.  - natural product research.  1909-1913. 

anisa s. al-hakimi, sarah abdul razak, n. omar, abdulwahab a. saeed, shamsul khamis, a. latiff.  (2022).  stem and leaf anatomical studies of selected species of barlerieae and ruellieae (acanthaceae) from yemen.  - sains malaysiana.  1609-1624. 

siti nurfaeiza abd razak, wan juliana wan ahmad, shamsul khamis, shukor md nor.  (2022).  morpho-physiological strategies of shorea leprosula miq. and shorea acuminata dyer in response to light intensity and nutrient amendments.  - forest.  1-18. 

muhamad faris osman, siti munirah mohd faudzi, shamsul khamis, shahrul razid sarbini, khozirah shaari.  (2022).  shedding light on pipers identity via computational mass spectrometry.  - malaysian journal of chemistry.  150-160. 

jumaat h. adam, mohd afiq aizat-juhari, nik nadira-farhana nik ariff, shamsul khamis, mohd hizamri mohd yasin, kiew-lian wan.  (2022).  rafflesia tunku-azizahiae (rafflesiaceae), a new species from pahang, malaysia.  - sains malaysiana.  3843-3855. 

ahmad azrul hakim yahaya, wan mohd nuzul hakimi wan salleh, nurunajah ab ghani, shamsul khamis, nurul syafiqah rezali, mohd afiq aizat juhari.  (2022).  chemical composition and bioactivities of magnolia candollii h.keng essential oil.  - zeitschrift fur naturforschung.  519-523. 

siti nur fatehah, r., shamsul, k., wan juliana, w. a., norela sulaiman, md. shahril,m. h., a. d. nurul darsani.  (2021).  distribution patterns of plant communities in riverine mangrove habitats and their associations with conductivity factors in the west coast of peninsular malaysia.  - the malaysian forester.  169-184. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, hakimi kassim, alene tawang.  (2021).  chemical composition of the essential oil of dysoxylum cauliflorum hiern (meliaceae).  - natural volatiles & essential oils.  7-12. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, hakimi kassim, alene tawang.  (2021).  essential oil composition of strychnos axillaris colebr. (loganiaceae).  - natural volatiles & essential oils.  13-17. 

a.k.n. hayati, k. shamsul, w.a.w. juliana, f.s. mohd. taib, m.n. shukor, m.h. shahril, e.a. filza.  (2021).  habitat and edaphic preference of rafflesia r. br. plant communities in royal belum state park, perak, peninsular malaysia.  - journal of environmental biology.  865-871. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2021).  composition of essential oil from actinodaphne sesquipedalis and its lipoxygenase activity.  - chemistry of natural compounds.  553-555. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2021).  chemical composition of the essential oil of diospyros argentea.  - chemistry of natural compounds.  556-557. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2021).  composition of volatile oil from anaxagorea javanica.  - chemistry of natural compounds.  558-559. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, muhammad nurakmal abdul rahman, mohd azlan nafiah.  (2021).  chemical composition and acetylcholinesterase activity of the essential oil of anisophyllea disticha.  - chemistry of natural compounds.  371-373. 

muhammad ammar mohd azhar, wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2021).  essential oil composition of beilschmiedia insignis from malaysia.  - chemistry of natural compounds.  374-375. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, salam ahmed abed.  (2021).  characterization of volatile components of chassalia chartacea and its acetylcholinesterase inhibitory activity.  - chemistry of natural compounds.  376-377. 

nor shafikah idris, muzzneena ahmad mustapha, norela sulaiman, shamsul khamis, shahril mohd husin, nurul darsani ahmad darbis.  (2021).  the dynamics of landscape changes surrounding a firefly ecotourism area.  - global ecology and conservation.  1-20. 

wan mohd nuzul hakimi wan salleh, mohd azlan nafiah, shamsul khamis, salam ahmed abed.  (2021).  chemical composition and lipoxygenase inhibitory activity of alseodaphne peduncularis meisn. essential oil.  - nat. volatiles & essent. oils.  29-33. 

rasyidah wahab, ahmad asnawi mus, kartini saibeh, shamsul khamis, handry mujih, geofarry gunsalam, dasini, ezron gerald, juhinin sariman, qarin aaron muhammad, qaid ahmad muhammad.  (2021).  soil physico-chemistry in the habitat of rafflesia in kinabalu park, sabah, malaysia..  - journal of tropical biology and conservation.  149-165. 

syarifah haniera sheikh kamal, mohd nazip suratman, shamsul khamis, ahmad najmi nik hassan, mohd syaiful mohammad.  (2021).  host-parasitic relationships between tetrastigma rafflesiae and rafflesia azlanii and rafflesia cantleyi in belum-temenggor forest complex, perak, malaysia.  - pertanika journal of tropical agricultural science.  755-771. 

muhamad faris osman, soo yee lee, shahrul razid sarbini, siti munirah mohd faudzi, shamsul khamis, badrul hisyam zainudin, khozirah shaari.  (2021).  metabolomics-driven discovery of an introduced species and two malaysian piper betle l. variants.  - plants.  1-19. 

shamsul, k., nik norafida, n. a., mat salleh, k. and a. latiff.  (2020).  tree species composition, diversity and biomass estimation in lower montane forest at fraser's hill, pahang, peninsular malaysia.  - the malaysian forester.  114-127. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2020).  chemical composition and lipoxygenase activity of the leaves essential oil of rothmannia macrophylla (hook.f.) bremek.  - journal of essential oil bearing plants.  331-336. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, muhammad nurakmal abdul rahman, mohd azlan nafiah.  (2020).  chemical composition and biological activities of dipterocarpus cornutus dyer essential oil.  - journal of biosciences: zeitschrift für naturforschung c.  171-175. 

a.k. nur hayati, k. shamsul, w.a. wan juliana, m. n. shukor, m. h. shahril, e. alyaa filza.  (2020).  plant community structure and diversity of the rafflesia habitat at the royal belum state park, perak, malaysia.  - the malaysian forester.  387-404. 

muhammad ammar mohd azhar, wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2020).  essential oil composition of three cryptocarya species from malaysia.  - zeitschrift fur naturforschung section c-a journal of biosciences.  297-301. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2020).  chemical composition of the essential oil of diospyros wallichii king & gamble (ebenaceae).  - natural volatiles & essential oils.  12-17. 

natasa mohd shakri, wan mohd nuzul hakimi wan salleh, shamsul khamis, nor azah mohamad ali, shazlyn milleana shaharudin.  (2020).  chemical composition of the essential oils of four polyalthia species from malaysia.  - zeitschrift fuer naturforschung, section c: a journal of biosciences.  473-478. 

natasa mohd shakri, wan mohd nuzul hakimi wan salleh, shamsul khamis, nor azah mohamad ali.  (2020).  chemical characterization of goniothalamus macrophyllus and goniothalamus malayanus leaves essential oils.  - zeitschrift fur naturforschung c a journal of biosciences.  485-488. 

natasa mohd shakri, wan mohd nuzul hakimi wan salleh, shamsul khamis, nor azah mohamad ali, muhammad helmi nadri.  (2020).  composition of the essential oils of three malaysian xylopia species (annonaceae).  - zeitschrift fur naturforschung c a journal of biosciences.  479-484. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2020).  chemical composition and anticholinesterase inhibitory activity of pavetta graciliflora wall. ex ridl. essential.  - zeitschrift fur naturforschung c a journal of biosciences.  467-471. 

wan mohd nuzul hakimi wan salleh, nur kamilah mohammed, nur athirah hashim, shamsul khamis, salam ahmed abed, saidu jibril.  (2020).  chemical constituents of beilschmiedia penangiana.  - chemistry of natural compounds.  576-577. 

sani aliyu, norazah basar1, yakubu rufai, shamsul khamis.  (2020).  phytochemical analysis and isolation of secondary metabolites from persea declinata.  - malaysian journal of analytical sciences.  970-977. 

faridah qz, ayuni m. n, shamsul khamis, anisa s. al-hakimi, rosimah nulit, norhidayah mad halid.  (2019).  traditional knowledge on aromatic plants as utilised by the jahai tribe in royal belum state park, perak, malaysia.  - malayan nature journal.  1-8. 

nur-athirah abdullah, siti nur fatehah radzi, lailatul-nadhirah asri, nor shafikah idris, shahril husin, azman sulaiman, shamsul khamis, norela sulaiman, izfa riza hazmi.  (2019).  insect community in riparian zones of sungai sepetang, sungai rembau and sungai chukai of peninsular malaysia.  - biodiversity data journal.  1-24. 

nur hayati a. k, shamsul k., wan juliana w. a., shahril m. h., alyaa filza e., mohamed shah redza h..  (2019).  re-assessment of the rafflesia populations and habitats in royal belum state park, perak, peninsular malaysia.  - malayan nature journal.  493-498. 

muhamad ikhwanuddin mat esa, rosimah nulit, shamsul khamis, rusea go.  (2019).  ethnomedical survey of plants used by the bateq in kuala koh, kelantan, malaysia.  - malayan nature journal.  439-451. 

shamsul k., m. s. nizam, anisa s. al-hakimi, n. talip.  (2019).  taxonomic value of petiole anatomy in malaysian litsea complex (lauraceae).  - malayan nature journal.  453-465. 

wan mohd nuzul hakimi wan salleh, nur athirah hashim, shamsul khamis.  (2019).  chemical constituents and lipoxygenase inhibitory activity of piper stylosum miq.  - bulletin of the chemical society of ethiopia.  587-592. 

wan mohd nuzul hakimi wan salleh, norkamilah abdullah, nur athirah hashim, heng yen khong, shamsul khamis.  (2019).  aporphine alkaloids from piper erecticaule and acetylcholinesterase inhibitory activity.  - boletin latinoamericano y del caribe de plantas medicinales y aromaticas.  527-532. 

nur athirah hashim, farediah ahmad, wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2019).  a new amide from piper maingayi hk.f. (piperaceae).  - natural product communications.  1-5. 

faridah qamaruz zaman, anisa s. al-hakimi, shamsul khamis, fatin f. ruhaizin and syuhada. m. zaidi.  (2018).  anatomy and histochemistry of structures producing aroma in leaves of sygyzium aromaticum (l.) merr. and clausena excavata burm. f..  - pertanika journal of tropical agricultural science.  .