muhammad izzelen izzauddin mamat, shamsul khamis, muhammad faqih luqman fuaad, nor zalipah mohamed, chik maslinda omar, daria mathew abdullah, wahizatul afzan azm.  (2023).  pollen sources foraged by domesticated stingless bee (heterotrigona itama) reared in gelam forests of terengganu, malaysia.  - serangga.  120-138. 

abubakar siddiq salihu, nurunajah ab ghani, nurulfazlina edayah rasol, shamsul khamis.  (2023).  chemical composition of the essential oil of litsea resinosa blume and acetylcholinesterase activity.  - journal of science and mathematics letters.  64-67. 

n.h.a. kadir, w.m.n.h.w. salleh, n.a. ghani, s. khamis, m.a.a juhairi.  (2023).  chemical composition and acetylcholinesterase activity of syzygium pyrifolium (blume) dc. essential oil.  - latin american applied research.  357-359. 

seemab akram, shamsul khamis, shahrizim zulkifly, rishzuan talib, nurul izza ab ghani.  (2023).  phylogeny study of 20 selected species of zingiberaceae from ex situ collections in peninsular malaysia.  - pertanika j. trop. agric. sci..  329-345. 

siti munirah mat yunoh, nur hayati abd. karim, halimah alias, mohd. noor mat isa, wan kiew lian, shamsul khamis, mohd afiq aizat juhari.  (2023).  rafflesia di pahang.  - .  21. 

wee suk ling, siti munirah mat yunoh, nur hayati abd. karim, shamsul khamis.  (2023).  rafflesia di pahang.  - .  22. 

mohd. hizamri mohd. yasin, norela sulaiman, shamsul khamis.  (2023).  rafflesia di pahang; rafflesia of pahang.  - .  1-69. 

seemab akram, shahrizim zulkifly, shamsul khamis, nurul izza ab ghani.  (2023).  preliminary in silico analysis of chs1 gene in commelinids clade: family zingiberaceae, costaceae, and poaceae.  - pertanika j. trop. agric. sci..  425-437. 

shamsul khamis mohd. afiq aizat juhari, siti nurfaeiza abd razak.  (2023).  hubungkait perumah rafflesia (tetrastigma sp.) dengan komuniti tumbuhan dan faktor tanah di hutan simpan jerantut tambahan, mukim ulu cheka, daerah jerantut, pahang.  - seminar ekspedisi saintifik kepelbagaian biologi hutan ulu cheka, jerantut, pahang..  . 

ahmad azrul hakim yahaya, wan mohd nuzul hakimi wan salleh, nurunajah ab ghani, shamsul khamis, nurul syafiqah rezali, mohd afiq aizat juhari.  (2022).  chemical composition and bioactivities of magnolia candollii h.keng essential oil.  - zeitschrift fur naturforschung.  519-523. 

jumaat h. adam, mohd afiq aizat-juhari, nik nadira-farhana nik ariff, shamsul khamis, mohd hizamri mohd yasin, kiew-lian wan.  (2022).  rafflesia tunku-azizahiae (rafflesiaceae), a new species from pahang, malaysia.  - sains malaysiana.  3843-3855. 

norela bt. sulaiman;siti norhafizah binti ahmad tarmidzi;mohd afiq aizat bin juhari;shamsul bin khamis.  (2022).  highland stingless bee identification and the assessment of domesticated stingless bees at high altitude.  - .  . 

muhamad faris osman, siti munirah mohd faudzi, shamsul khamis, shahrul razid sarbini, khozirah shaari.  (2022).  shedding light on pipers identity via computational mass spectrometry.  - malaysian journal of chemistry.  150-160. 

nur-fatehah sr, nur-shafikah i, shamsul k, wan-juliana wa, norela sulaiman, shahril mh, nurul-darsani ad.  (2022).  stand structure and productivity of the firefly riverine mangrove habitat at three rivers in peninsular malaysia.  - journal of tropical forest science.  114-126. 

muzzneena binti ahmad mustapha;norela bt. sulaiman;shamsul bin khamis.  (2022).  mapping potential firefly habitats using texture based classification of satellite imagery.  - .  . 

siti nurfaeiza abd razak, wan juliana wan ahmad, shamsul khamis, shukor md nor.  (2022).  morpho-physiological strategies of shorea leprosula miq. and shorea acuminata dyer in response to light intensity and nutrient amendments.  - forest.  1-18. 

syarifah haniera sheikh kamal, mohd nazip suratman, shamsul khamis, ahmad najmi nik hassan, mohd syaiful mohammad.  (2022).  growth rate, mortality rate and life cycle of rafflesia azlanii and r. cantleyi in belum-temenggor forest complex, perak, malaysia.  - sains malaysiana.  943-957. 

a. r. siti nurfaeiza, k. shamsul, w.a. wan juliana, m.n. shukor.  (2022).  tropical rainforest regeneration in an area devastated by dam impoundment in hulu terengganu, malaysia.  - the malaysian forester.  96-108. 

shamsul khamis, noralasikin yahya, rukayah aman.  (2022).  kamus herba fitoterapi.  - kamus fitoterapi.  1-61. 

anisa s. al-hakimi, sarah abdul razak, n. omar, abdulwahab a. saeed, shamsul khamis, a. latiff.  (2022).  stem and leaf anatomical studies of selected species of barlerieae and ruellieae (acanthaceae) from yemen.  - sains malaysiana.  1609-1624. 

wan mohd nuzul hakimi wan salleh, natasa mohd shakri, shamsul khamis, william n. setzer, muhammad helmi nadri.  (2022).  chemical composition of three malaysian horsfieldia essential oils.  - natural product research.  1909-1913. 

m.a.m. azhar, w.m.n.h.w. salleh, s. khamis, n.a. ghani.  (2022).  variation in essential oil composition of three litsea species from malaysia.  - rivista italiana delle sostanze grasse.  57-61. 

w.m.n.h.w. salleh, n.m. shakri, m.a. nafiah, s. khamis.  (2022).  chemical constituents and acetylcholinesterase activity from polyalthia sumatrana (miq.) kurz.  - latin american applied research.  73-76. 

s akram, n i ab ghani1,, s khamis and s zulkifly.  (2021).  newly designed chs genic primers for four zingiberaceae species (alpinia mutica, alpinia rafflesiana, hornstedtia leonurus and scaphochlamys kunstleri).  - iop conference series: earth and environmental science.  1-9. 

a.k.n. hayati, k. shamsul, w.a.w. juliana, f.s. mohd. taib, m.n. shukor, m.h. shahril, e.a. filza.  (2021).  habitat and edaphic preference of rafflesia r. br. plant communities in royal belum state park, perak, peninsular malaysia.  - journal of environmental biology.  865-871. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2021).  composition of essential oil from actinodaphne sesquipedalis and its lipoxygenase activity.  - chemistry of natural compounds.  553-555. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2021).  chemical composition of the essential oil of diospyros argentea.  - chemistry of natural compounds.  556-557. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2021).  composition of volatile oil from anaxagorea javanica.  - chemistry of natural compounds.  558-559. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, muhammad nurakmal abdul rahman, mohd azlan nafiah.  (2021).  chemical composition and acetylcholinesterase activity of the essential oil of anisophyllea disticha.  - chemistry of natural compounds.  371-373. 

muhammad ammar mohd azhar, wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2021).  essential oil composition of beilschmiedia insignis from malaysia.  - chemistry of natural compounds.  374-375. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, salam ahmed abed.  (2021).  characterization of volatile components of chassalia chartacea and its acetylcholinesterase inhibitory activity.  - chemistry of natural compounds.  376-377. 

nor shafikah idris, muzzneena ahmad mustapha, norela sulaiman, shamsul khamis, shahril mohd husin, nurul darsani ahmad darbis.  (2021).  the dynamics of landscape changes surrounding a firefly ecotourism area.  - global ecology and conservation.  1-20. 

wan mohd nuzul hakimi wan salleh, mohd azlan nafiah, shamsul khamis, salam ahmed abed.  (2021).  chemical composition and lipoxygenase inhibitory activity of alseodaphne peduncularis meisn. essential oil.  - nat. volatiles & essent. oils.  29-33. 

siti nur fatehah, r., shamsul, k., wan juliana, w. a., norela sulaiman, md. shahril,m. h., a. d. nurul darsani.  (2021).  distribution patterns of plant communities in riverine mangrove habitats and their associations with conductivity factors in the west coast of peninsular malaysia.  - the malaysian forester.  169-184. 

rasyidah wahab, ahmad asnawi mus, kartini saibeh, shamsul khamis, handry mujih, geofarry gunsalam, dasini, ezron gerald, juhinin sariman, qarin aaron muhammad, qaid ahmad muhammad.  (2021).  soil physico-chemistry in the habitat of rafflesia in kinabalu park, sabah, malaysia..  - journal of tropical biology and conservation.  149-165. 

syarifah haniera sheikh kamal, mohd nazip suratman, shamsul khamis, ahmad najmi nik hassan, mohd syaiful mohammad.  (2021).  host-parasitic relationships between tetrastigma rafflesiae and rafflesia azlanii and rafflesia cantleyi in belum-temenggor forest complex, perak, malaysia.  - pertanika journal of tropical agricultural science.  755-771. 

nor akmalazura jani, farah aina rokman, rashidah iberahim, shamsul khamis.  (2021).  composition and bioactivities of the rhizomes essential oil of curcuma aeruginosa.  - gading journal of science and technology.  66-74. 

muhamad faris osman, soo yee lee, shahrul razid sarbini, siti munirah mohd faudzi, shamsul khamis, badrul hisyam zainudin, khozirah shaari.  (2021).  metabolomics-driven discovery of an introduced species and two malaysian piper betle l. variants.  - plants.  1-19. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, hakimi kassim, alene tawang.  (2021).  chemical composition of the essential oil of dysoxylum cauliflorum hiern (meliaceae).  - natural volatiles & essential oils.  7-12. 

shamsul khamis, nur syamina jamaluddin.  (2021).  an annotated checklist of flowering plants from foothill to seroja camp trail in mount korbu perak semenanjung malaysia.  - seminar saintifik geobiotrail gunung korbu 2021.  183-188. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, hakimi kassim, alene tawang.  (2021).  essential oil composition of strychnos axillaris colebr. (loganiaceae).  - natural volatiles & essential oils.  13-17. 

sani aliyu, norazah basar1, yakubu rufai, shamsul khamis.  (2020).  phytochemical analysis and isolation of secondary metabolites from persea declinata.  - malaysian journal of analytical sciences.  970-977. 

syarifah haniera sheikh kamal, mohd nazip suratman, shamsul khamis, ahmad najmi nik hassan, mohd syaiful mohammad.  (2020).  host-parasitic relationships between tetrastigma rafflesiae and rafflesia azlanii and r. cantleyi in belum-temenggor forest complex, perak.  - 7th international conference on science and social research 2020.  1-19. 

wahab, rasyidah, saibeh, k., s. khamis,.  (2020).  tekstur tanah di habitat bunga rafflesia sp. di kawasan taman negara kinabalu, sabah, malaysia.  - the 12th postgraduate symposium,.  20. 

nur syamina jamaluddin, shamsul khamis.  (2020).  a collection of sapotaceae in fraser hill, pahang, peninsular malaysia.  - fraser hill symposium.  1-18. 

shamsul, k., siti nur fateha radzi, siti nur syazwani musthami@sohaimi, nur syamina jamaludin, radhiah zakaria, ahmad fitri, z., mohd nizam, m.s., talip, n., latiff, a..  (2020).  hutan ulu sat: nadi pemeliharaan alam semulajadi kelantan (ulu sat forest: the heart of kelantan's nature conservation).  - .  13. 

natasa mohd shakri, wan mohd nuzul hakimi wan salleh, shamsul khamis, nor azah mohamad ali, shazlyn milleana shaharudin.  (2020).  chemical composition of the essential oils of four polyalthia species from malaysia.  - zeitschrift fuer naturforschung, section c: a journal of biosciences.  473-478. 

norela sulaiman, azman sulaiman, muzzneena ahmad mustapha, izfa riza hazmi, farah shafawati mohd taib, saiful mansor, shamsul khamis, abdullah samat, juliana senawi, salmah yaakop, nurul wahida othman, hani kartini agustar,.  (2020).  majalah knovasi 1/2020.  - pembelajaran aktif di lapangan untuk pemuliharaan kelip-kelip.  1. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2020).  chemical composition of the essential oil of diospyros wallichii king & gamble (ebenaceae).  - natural volatiles & essential oils.  12-17. 

shamsul, k., nik norafida, n. a., mat salleh, k. and a. latiff.  (2020).  tree species composition, diversity and biomass estimation in lower montane forest at fraser's hill, pahang, peninsular malaysia.  - the malaysian forester.  114-127. 

muhammad ammar mohd azhar, wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2020).  essential oil composition of three cryptocarya species from malaysia.  - zeitschrift fur naturforschung section c-a journal of biosciences.  297-301. 

a.k. nur hayati, k. shamsul, w.a. wan juliana, m. n. shukor, m. h. shahril, e. alyaa filza.  (2020).  plant community structure and diversity of the rafflesia habitat at the royal belum state park, perak, malaysia.  - the malaysian forester.  387-404. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, muhammad nurakmal abdul rahman, mohd azlan nafiah.  (2020).  chemical composition and biological activities of dipterocarpus cornutus dyer essential oil.  - journal of biosciences: zeitschrift für naturforschung c.  171-175. 

shamsul khamis, wan mohd nazri abdul rahman, aiza harun.  (2020).  63 tumbuhan ubatan.  - .  136. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2020).  chemical composition and lipoxygenase activity of the leaves essential oil of rothmannia macrophylla (hook.f.) bremek.  - journal of essential oil bearing plants.  331-336. 

norela sulaiman, muzzneena ahmad mustapha, salmah yaakop, azman sulaiman, izfa riza hazmi, nurul wahida othman, juliana senawi, farah shafawati mohd taib, saiful mansor, abdullah samat, hani kartini agustar, shamsul khamis, lailatul nadhirah asri, nur ati.  (2020).  kelip-kelip anugerah alam di waktu malam.  - .  30. 

k. shamsul, a. norhayati, y.f. ng, a. latiff.  (2020).  biodiversity of water catchment forest: hutan simpan bukit cherakah, selangor.  - .  31. 

k. shamsul, a. norhayati, y.f. ng, a. latiff.  (2020).  biodiversity of water catchment forest: hutan simpan bukit cherakah, selangor..  - .  25. 

k. shamsul, a. norhayati, y.f. ng, a. latiff.  (2020).  biodiversity of water catchment forest: hutan simpan bukit cherakah, selangor.  - .  12. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2020).  chemical composition and anticholinesterase inhibitory activity of pavetta graciliflora wall. ex ridl. essential.  - zeitschrift fur naturforschung c a journal of biosciences.  467-471. 

natasa mohd shakri, wan mohd nuzul hakimi wan salleh, shamsul khamis, nor azah mohamad ali, muhammad helmi nadri.  (2020).  composition of the essential oils of three malaysian xylopia species (annonaceae).  - zeitschrift fur naturforschung c a journal of biosciences.  479-484. 

natasa mohd shakri, wan mohd nuzul hakimi wan salleh, shamsul khamis, nor azah mohamad ali.  (2020).  chemical characterization of goniothalamus macrophyllus and goniothalamus malayanus leaves essential oils.  - zeitschrift fur naturforschung c a journal of biosciences.  485-488. 

wan mohd nuzul hakimi wan salleh, nur kamilah mohammed, nur athirah hashim, shamsul khamis, salam ahmed abed, saidu jibril.  (2020).  chemical constituents of beilschmiedia penangiana.  - chemistry of natural compounds.  576-577. 

shamsul khamis, kushairi salleh mohd.  (2020).  bukit fraser kepelbagaian biology dan persekitaran fizikal.  - .  13. 

shamsul bin khamis;mohd. nizam bin mohd. said.  (2020).  kajian taksonomi dan ekologi ke atas famili lauraceae di semenanjung malaysia.  - .  . 

muhamad ikhwanuddin mat esa, rosimah nulit, shamsul khamis, rusea go.  (2019).  ethnomedical survey of plants used by the bateq in kuala koh, kelantan, malaysia.  - malayan nature journal.  439-451. 

shamsul k., m. s. nizam, anisa s. al-hakimi, n. talip.  (2019).  taxonomic value of petiole anatomy in malaysian litsea complex (lauraceae).  - malayan nature journal.  453-465. 

wan mohd nuzul hakimi wan salleh, nur athirah hashim, shamsul khamis.  (2019).  chemical constituents and lipoxygenase inhibitory activity of piper stylosum miq.  - bulletin of the chemical society of ethiopia.  587-592. 

wan mohd nuzul hakimi wan salleh, norkamilah abdullah, nur athirah hashim, heng yen khong, shamsul khamis.  (2019).  aporphine alkaloids from piper erecticaule and acetylcholinesterase inhibitory activity.  - boletin latinoamericano y del caribe de plantas medicinales y aromaticas.  527-532. 

nur athirah hashim, farediah ahmad, wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2019).  a new amide from piper maingayi hk.f. (piperaceae).  - natural product communications.  1-5. 

mohd lip jabit, mohd nazrul hisham daud, shamsul khamis.  (2019).  cytotoxic, nitric oxide and alpha-glucosidase inhibitory activities of biflavonoids from garcinia prainiana king.  - advances in applied chemistry and biochemistry.  55-67. 

r siti nur fatehah, k shamsul, w a wan juliana norela sulaiman, m h md. shahril, a d nurul darsani, m s aisah.  (2019).  riverine mangrove plant species composition and vegetation structure of the fireflys habitat at sungai sepetang kampung dew, taiping, perak.  - 14th international annual symposium universiti malaysia terengganu.  47. 

nur hayati abd karim, shamsul khamis, wan juliana wan ahmad.  (2019).  reassessment of the rafflesia population and microhabitat in royal belum state park, perak.  - international conference on tropical forest science (ictfs): forest for sustainable development goals.  32. 

nur-athirah abdullah, siti nur fatehah radzi, lailatul-nadhirah asri, nor shafikah idris, shahril husin, azman sulaiman, shamsul khamis, norela sulaiman, izfa riza hazmi.  (2019).  insect community in riparian zones of sungai sepetang, sungai rembau and sungai chukai of peninsular malaysia.  - biodiversity data journal.  1-24. 

nur hayati a. k, shamsul k., wan juliana w. a., shahril m. h., alyaa filza e., mohamed shah redza h..  (2019).  re-assessment of the rafflesia populations and habitats in royal belum state park, perak, peninsular malaysia.  - malayan nature journal.  493-498. 

shamsul k., nizam, m.s., al-hakimi s. anisa, talip n..  (2019).  taxonomic value of petiole anatomy in malaysian litsea complex (lauraceae).  - 11th flora malesiana symposium.  110. 

shaharuddin mohamad ismail, shamsul khamis, nurul hidayah shamsul arrifin, faizal mohamad hassan mukhtar.  (2019).  penjelajahan flora nur laman bestari eco resort.  - .  127. 

mazlan mohd som, shahril mod husin, aisyah md shukor, alyaa filza effendi, noor ilyani abd rani, shamsul khamis, nur hayati abdul karim, shukor md. nor.  (2019).  rafflesia conservation efforts and management at royal belum state park.  - .  149. 

faridah qz, ayuni m. n, shamsul khamis, anisa s. al-hakimi, rosimah nulit, norhidayah mad halid.  (2019).  traditional knowledge on aromatic plants as utilised by the jahai tribe in royal belum state park, perak, malaysia.  - malayan nature journal.  1-8. 

nurul ayuni, m.n., faridah, q.z., anisa, s.a., rosimah, n., norhidayah, m.h. & shamsul, k..  (2018).  ethnobotanical documentation of plants used by the jahai tribe in royal belum state park, perak.  - proceedings of the 15th seminar on medicinal and aromatic plants (maps-15).  . 

nik norhazrina nik mohd kamil, ahmad fitri zahari, shamsul khamis, nur aqilah mustafa bakray, haja maideen kader maiden.  (2018).  a natural heritage: the flora and fauna of universiti kebangsaan malaysia.  - .  8. 

norela sulaiman, shamsul khamis, azman sulaiman & muzzneena ahmad mustapha.  (2018).  penanaman berembang: usaha pemuliharaan kelip-kelip.  - rampai penyelidikan siri 6. stem untuk komuniti.  . 

rasyidah wahab, kartini saibeh, shamsul khamis & monica suleiman.  (2018).  kajian komuniti tumbuhan bagi perumah rafflesia (tetrastigma sp.) di lokasi berbeza di taman negara kinabalu, sabah.  - .  . 

ahmad fariz bin mohamed;riza bin sulaiman;shaharudin bin idrus;bisharuzi bin omar;khairul nizam bin abdul maulud;nik mohd noor faizul bin md saad;nor`ain binti mohd jali;rd puteri khairani binti khirotdin;shamsul bin khamis.  (2018).  ukm campus tree id and hazardous trees assessment.  - .  . 

faridah qamaruz zaman, anisa s. al-hakimi, shamsul khamis, fatin f. ruhaizin and syuhada. m. zaidi.  (2018).  anatomy and histochemistry of structures producing aroma in leaves of sygyzium aromaticum (l.) merr. and clausena excavata burm. f..  - pertanika journal of tropical agricultural science.  . 

soo foong lian, shamsulbin khamis & noraini talip.  (2017).  comparative plant morphology and leaf anatomy of selected accessions of ipomoea batatas (l.) lam. in peninsular malaysia.  - fundamental science congress 2017.  .