khamis, shamsul, mat yunoh, siti-munirah.  (2025).  rafflesia of peninsular malaysia: research and conservation efforts..  - 10th european botanic garden congress.  . 

gregorius richard clay rudyson, siti zaiton mat so’ad, elok zubaidah, shamsul khamis .  (2025).  a phytochemical profiling and in vitro antimicrobial evaluation of methanolic extract and fractions of dicranopteris linearis leaves.  - journal of pharmacy .  305 - 315. 

shamsul khamis, abdul latif mohamad, norhayati ahmad & juliana senawi.  (2025).  laporan akhir: pelan induk untuk penyelidikan, pendidikan awam & pembangunan ekopelancongan di pusat pendidikan alam dan penyelidikan, endau rompin, johor darul takzim.  - .  . 

siti noor syafarehan, m.i. , nurul azira, m.z. , shamsul, k. hamdan, o., norhayati, a.  (2025).  taburan flora pada mikrohabitat berbeza di bukit gua madu, kelantan.  - fakult sains dan teknologi, ukm .  . 

nurul azira, m.z., siti noor syafarehan, m.i., shamsul, k. dan rafidah, a.r .  (2025).  komposisi flora hutan batu kapur terhadap mikrohabitat di bukit kurau,perak dan gua ikan kelantan.  - simposium biologi kebangsaan 2025.  . 

shamsul khamis, siti nurfaeiza abd razak, ras elmeera mohd. rashid, siti noor syafarehan mohd. isa, a. latiff, nadzifah yaakub, hafizan juahir, norhayati ngah, mohd. nordin abdul rahman, mohd. khairul amri kamarudin, jeyanny vijayanathan.  (2025).  taman rimba ilmu tanah bris (tribe), universiti sultan zainal abidin kampus besut, terengganu, peninsular malaysia.  - proceedings of the ist botanic gardens symposium 2025.  54 - 58. 

najwa oma, norazian mohd hassan, shamsul khamis.  (2025).  screening of phytochemicals, determination of total phenolic and flavonoid contents, and antioxidant characteristics of leaves of guioa diplopetala (hassk.) radlk.  - proceedings 1st botanic gardens symposium 2025.  99 - 103. 

nur hanna insyirah mohd rahim, shamsul khamis.  (2025).  morphological description of garcinia sp. (clusiaceae) from fraser’s hill, pahang.  - undergraduate research journal for biological sciences and biotechnology .  88 - 95. 

nursabrina najwa salmin, nurulfazlina edayah rasol, adiana mohamed adib, fauziah abdullah, ling sui kiong, shamsul khamis, nor hadiani ismail.  (2025).  comprehensive phytochemical profiling of malaysian piper sarmentosum leaves using mass-based dereplication and molecular networking.  - malaysian journal of chemistry.  36 - 52. 

siti nurfaeiza abd razak, shamsul khamis, mohd afiq aizat juhari.  (2025).  hubung kait perumah rafflesia (tetrastigma sp.) dengan komuniti tumbuhan dan ciri mikrohabitat di hutan simpan jerantut tambahan, mukim ulu cheka, daerah jerantut, pahang.  - sains malaysiana.  677 - 689. 

siti noor syafarehan mohd. isa, nurul azira mohd. zamziba, shamsul khamis, hamdan omar & norhayati ahmad .  (2025).  kepelbagaian flora pada mikrohabitat berbeza di bukit lang, ipoh, perak.  - hari penyelidikan dan inovasi (hapi) 2025.  . 

nurul azira mohd zamziba, siti noor syafarehan mohd isa, shamsul khamis, rafidah abdul rahman.  (2025).  kepelbagaian flora pada mikrohabitat berbeza di bukit kurau, perak.  - hari penyelidikan dan inovasi (hapi) 2025.  . 

muhamad faris osman, siti munirah mohd faudzi, khozirah shaari, shahrul razid sarbini, shamsul khamis.  (2025).  recognition of sesquiterpenoids and piperidine alkamides as two discerning metabolite classes in the fruits of *piper nigrum* 'semongok aman'.  - borneo journal of resource science and technology.  130 - 145. 

shah badri mohd nor, ahmad fariz mohamed, shamsul khamis.  (2025).  forest certification management experience in asia pacific and europe: a case study of malaysia and sweden.  - journal of forest science.  269 - 285. 

shamsul khamis, siti nurfaeiza abd razak, ras elmeera mohd rashid, loh khai vee, nadzifah yaakub, hafizan juahir, mohd nordin abdul rahman, jeyanny vijayanathan.  (2025).  tree species composition and community structure of vegetation on bris soil of coastal terengganu, peninsular malaysia.  - the malayan nature journal.  125 - 134. 

milan javier maidm, shamsul khamis.  (2024).  kepelbagaian tumbuhan herba terestrial di bukit fraser, pahang (diversity of terrestrial herbaceous plants in fraser’s hill, pahang).  - semarak proceedings of natural and environmental science.  12 - 17. 

shamsul khamis, haja maideen kader maideen, nur aliah mohamad khaduwi, siti noor syafarehan mohd. isa.  (2024).  kepelbagaian flora taman botani negara shah alam, selangor.  - .  . 

ras elmeera mohd rashid, shamsul khamis.  (2024).  tree species composition and forest structure in natural vegetation of bris soil on coastal of terengganu, east peninsular malaysia .  - prosiding simposium prasiswazah fst 2024 gugusan sains biologi dan bioteknologi.  125 - 131. 

khamis shamsul, siti nur hannah ismail, mohamad mansor, ithnin irdazulfa, ali mohd saifuddin, buang sabarudin, sajat azmiah .  (2024).  enhancing ecosystem services and urban sustainability through tree diversity in malaysian street landscapes .  - international urban tree diversity conference.  74 - 75. 

roberta chaya tawie tingga , millawati gani , nur azimah osman , nor rahman aifat , eddie chan , shamsul khamis , emelda rosseleena rohani , norlinda mohd-daut , abd rahman mohd-ridwan, badrul munir md-zain.  (2024).  interspecific variation in the diet of symphalangus syndactylus and macaca nemestrina at genting highlands, pahang, peninsular malaysia.  - biodiversity data journal.  1 - 18. 

haja maideen, norlaila najwa rahman, nur aliah, nurul nadhirah, noraini talip, nik norhazrina, nur aqilah bakray, ahmad fitri zohari, nur farah ain zainee, mohd nizam mohd said, shamsul khamis, nur syazwani mohamad basir.  (2024).  anatomical characteristics of selected dennstaedtiaceae species from peninsular malaysia.  - malaysian applied biology.  229 - 237. 

shamsul khamis1,2, siti nurfaeiza abd razak3, ras elmeera mohd rashid2, siti noor syafarehan mohd isa2, abdul latiff mohamad2, nadzifah yaakub4, hafizan juahir4, norhayati ngah4, mohd nordin abdul rahman5, mohd khairul amri kamaruddin5 & jeyanny vijayana.  (2024).  diversity and potential of aromatic plants in taman rimba ilmu tanah bris (tribe) unisza campus besut, terengganu, east peninsular malaysia..  - 16th medicinal and aromatic plants seminar (maps-16), 2024.  1-43. 

shamsul khamis, ras elmeera mohd rashid, siti noor syafarehan mohd isa, abdul latiff mohamad, siti nurfaeiza abd razak, nadzifah yaakub, hafizan juahir, norhayati ngah, mohd nordin abdul rahman, mohd khairul amri kamaruddin, jeyanny vijayan.  (2024).  a preliminary checklist of flowering plants in taman rimba ilmu tanah bris (tribe) unisza kampus besut, terengganu, east peninsular malaysia..  - botanic gardens symposium 2024.  1-2. 

roberta chaya tawie tingga, millawati gani, nur azimah osman, nor rahman aifat, eddie chan, shamsul khamis, emelda rosseleena rohani, norlinda mohd-daut, abd rahman mohd ridwan, badrul munir md-zain.  (2024).  interspecific variation in the diet of symphalangus syndactylus and macaca nemestrina at genting highlands, pahang, peninsular malaysia.  - biodiversity data journal.  1-18. 

noraini talip, jayasrie eyamalay, mohamad ruzi abdul rahman, mohd nizam mohd said, haja maideen kader maideen, nik norhazrina nik mohd kamil, nur aqilah mustafa bakray, shamsul khamis, ahmad fitri zohari, nur farah ain zainee, hamidun bunawan.  (2024).  ciri adaptasi anatomi daun bougainvillea spectabilis w. yang dijangkiti virus bougainvillea chlorotic vein-banding.  - malaysian applied biology.  43-54. 

wan mohd nuzul hakimi wan salleh, mohd hafiz arzmi, shamsul khamis, mohd afiq aizat juhari.  (2024).  chemical composition and antioxidant activity of the essential oil of tabernaemontana dichotoma.  - chemistry of natural compounds.  169-170. 

nurunajah ab ghani, wan mohd nuzul hakimi wan salleh, abubakar siddiq salihu, nurulfazlina edayah rasol, shamsul khamis.  (2024).  essential oil composition of antidesma cuspidatum and acetylcholinesterase inhibitory activity.  - chemistry of natural compounds.  167-168. 

khamis shamsul, ismail siti nur hannah ismail, mohamad mansor, simis mazifah, ali mohd saifuddin, ithnin irda zulfa, mahmud khairil, buang sabarudin, sajat azmiah.  (2024).  enhancing ecosystem services through urban tree diversity in malaysian urban street landscapes.  - urban tree diversity 5, 2024.  . 

shamsul khamis.  (2024).  plant community and edaphic factors in selected rafflesia habitats in malaysia.  - kelantan international congress on rafflesia 2024 (kicr2024) .  . 

sofia castelo, jamie a. carr, shamsul khamis, melissa sivaraj, tainan messina.  (2024).  a study of climate resilience in urban tree species of peninsular malaysia.  - eco cities.  1 - 18. 

seemab akram, shamsul khamis, shahrizim zulkifly, rishzuan talib, nurul izza ab ghani.  (2023).  phylogeny study of 20 selected species of zingiberaceae from ex situ collections in peninsular malaysia.  - pertanika j. trop. agric. sci..  329-345. 

abubakar siddiq salihu, nurunajah ab ghani, nurulfazlina edayah rasol, shamsul khamis.  (2023).  chemical composition of the essential oil of litsea resinosa blume and acetylcholinesterase activity.  - journal of science and mathematics letters.  64-67. 

m e aznan, z a rahman, s n a tarmidzi, w m r idris, t lihan, s khamis, a a kadir, n a a jalil, m r.  (2023).  preliminary soil characteristics and growth performance of selected shrub plants for bio-engineering technique.  - international cenference on science and technology application in climate change (staclim-2022).  1-11. 

seemab akram, shahrizim zulkifly, shamsul khamis, nurul izza ab ghani.  (2023).  preliminary in silico analysis of chs1 gene in commelinids clade: family zingiberaceae, costaceae, and poaceae.  - pertanika j. trop. agric. sci..  425-437. 

n.h.a. kadir, w.m.n.h.w. salleh, n.a. ghani, s. khamis, m.a.a juhairi.  (2023).  chemical composition and acetylcholinesterase activity of syzygium pyrifolium (blume) dc. essential oil.  - latin american applied research.  357-359. 

muhammad izzelen izzauddin mamat, shamsul khamis, muhammad faqih luqman fuaad, nor zalipah mohamed, chik maslinda omar, daria mathew abdullah, wahizatul afzan azm.  (2023).  pollen sources foraged by domesticated stingless bee (heterotrigona itama) reared in gelam forests of terengganu, malaysia.  - serangga.  120-138. 

shamsul khamis mohd. afiq aizat juhari, siti nurfaeiza abd razak.  (2023).  hubungkait perumah rafflesia (tetrastigma sp.) dengan komuniti tumbuhan dan faktor tanah di hutan simpan jerantut tambahan, mukim ulu cheka, daerah jerantut, pahang.  - seminar ekspedisi saintifik kepelbagaian biologi hutan ulu cheka, jerantut, pahang..  . 

mohd. hizamri mohd. yasin, norela sulaiman, shamsul khamis.  (2023).  rafflesia di pahang; rafflesia of pahang.  - .  1-69. 

wee suk ling, siti munirah mat yunoh, nur hayati abd. karim, shamsul khamis.  (2023).  pengenalan kepada rafflesia di malaysia.  - .  16-37. 

siti munirah mat yunoh, nur hayati abd. karim, halimah alias, mohd. noor mat isa, wan kiew lian, shamsul khamis, mohd afiq aizat juhari.  (2023).  pengecaman spesies rafflesia.  - .  48-69. 

a. r. siti nurfaeiza, k. shamsul, w.a. wan juliana, m.n. shukor.  (2022).  tropical rainforest regeneration in an area devastated by dam impoundment in hulu terengganu, malaysia.  - the malaysian forester.  96-108. 

norela bt. sulaiman;siti norhafizah binti ahmad tarmidzi;mohd afiq aizat bin juhari;shamsul bin khamis.  (2022).  highland stingless bee identification and the assessment of domesticated stingless bees at high altitude.  - .  . 

nur-fatehah sr, nur-shafikah i, shamsul k, wan-juliana wa, norela sulaiman, shahril mh, nurul-darsani ad.  (2022).  stand structure and productivity of the firefly riverine mangrove habitat at three rivers in peninsular malaysia.  - journal of tropical forest science.  114-126. 

syarifah haniera sheikh kamal, mohd nazip suratman, shamsul khamis, ahmad najmi nik hassan, mohd syaiful mohammad.  (2022).  growth rate, mortality rate and life cycle of rafflesia azlanii and r. cantleyi in belum-temenggor forest complex, perak, malaysia.  - sains malaysiana.  943-957. 

w.m.n.h.w. salleh, n.m. shakri, m.a. nafiah, s. khamis.  (2022).  chemical constituents and acetylcholinesterase activity from polyalthia sumatrana (miq.) kurz.  - latin american applied research.  73-76. 

m.a.m. azhar, w.m.n.h.w. salleh, s. khamis, n.a. ghani.  (2022).  variation in essential oil composition of three litsea species from malaysia.  - rivista italiana delle sostanze grasse.  57-61. 

wan mohd nuzul hakimi wan salleh, natasa mohd shakri, shamsul khamis, william n. setzer, muhammad helmi nadri.  (2022).  chemical composition of three malaysian horsfieldia essential oils.  - natural product research.  1909-1913. 

anisa s. al-hakimi, sarah abdul razak, n. omar, abdulwahab a. saeed, shamsul khamis, a. latiff.  (2022).  stem and leaf anatomical studies of selected species of barlerieae and ruellieae (acanthaceae) from yemen.  - sains malaysiana.  1609-1624. 

shamsul khamis, noralasikin yahya, rukayah aman.  (2022).  kamus herba fitoterapi.  - .  1-61. 

muzzneena binti ahmad mustapha;norela bt. sulaiman;shamsul bin khamis.  (2022).  mapping potential firefly habitats using texture based classification of satellite imagery.  - .  . 

siti nurfaeiza abd razak, wan juliana wan ahmad, shamsul khamis, shukor md nor.  (2022).  morpho-physiological strategies of shorea leprosula miq. and shorea acuminata dyer in response to light intensity and nutrient amendments.  - forest.  1-18. 

muhamad faris osman, siti munirah mohd faudzi, shamsul khamis, shahrul razid sarbini, khozirah shaari.  (2022).  shedding light on pipers identity via computational mass spectrometry.  - malaysian journal of chemistry.  150-160. 

jumaat h. adam, mohd afiq aizat-juhari, nik nadira-farhana nik ariff, shamsul khamis, mohd hizamri mohd yasin, kiew-lian wan.  (2022).  rafflesia tunku-azizahiae (rafflesiaceae), a new species from pahang, malaysia.  - sains malaysiana.  3843-3855. 

ahmad azrul hakim yahaya, wan mohd nuzul hakimi wan salleh, nurunajah ab ghani, shamsul khamis, nurul syafiqah rezali, mohd afiq aizat juhari.  (2022).  chemical composition and bioactivities of magnolia candollii h.keng essential oil.  - zeitschrift fur naturforschung.  519-523. 

siti nur fatehah, r., shamsul, k., wan juliana, w. a., norela sulaiman, md. shahril,m. h., a. d. nurul darsani.  (2021).  distribution patterns of plant communities in riverine mangrove habitats and their associations with conductivity factors in the west coast of peninsular malaysia.  - the malaysian forester.  169-184. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, hakimi kassim, alene tawang.  (2021).  chemical composition of the essential oil of dysoxylum cauliflorum hiern (meliaceae).  - natural volatiles & essential oils.  7-12. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, hakimi kassim, alene tawang.  (2021).  essential oil composition of strychnos axillaris colebr. (loganiaceae).  - natural volatiles & essential oils.  13-17. 

a.k.n. hayati, k. shamsul, w.a.w. juliana, f.s. mohd. taib, m.n. shukor, m.h. shahril, e.a. filza.  (2021).  habitat and edaphic preference of rafflesia r. br. plant communities in royal belum state park, perak, peninsular malaysia.  - journal of environmental biology.  865-871. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2021).  composition of essential oil from actinodaphne sesquipedalis and its lipoxygenase activity.  - chemistry of natural compounds.  553-555. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2021).  chemical composition of the essential oil of diospyros argentea.  - chemistry of natural compounds.  556-557. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2021).  composition of volatile oil from anaxagorea javanica.  - chemistry of natural compounds.  558-559. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, muhammad nurakmal abdul rahman, mohd azlan nafiah.  (2021).  chemical composition and acetylcholinesterase activity of the essential oil of anisophyllea disticha.  - chemistry of natural compounds.  371-373. 

muhammad ammar mohd azhar, wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2021).  essential oil composition of beilschmiedia insignis from malaysia.  - chemistry of natural compounds.  374-375. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, salam ahmed abed.  (2021).  characterization of volatile components of chassalia chartacea and its acetylcholinesterase inhibitory activity.  - chemistry of natural compounds.  376-377. 

nor shafikah idris, muzzneena ahmad mustapha, norela sulaiman, shamsul khamis, shahril mohd husin, nurul darsani ahmad darbis.  (2021).  the dynamics of landscape changes surrounding a firefly ecotourism area.  - global ecology and conservation.  1-20. 

wan mohd nuzul hakimi wan salleh, mohd azlan nafiah, shamsul khamis, salam ahmed abed.  (2021).  chemical composition and lipoxygenase inhibitory activity of alseodaphne peduncularis meisn. essential oil.  - nat. volatiles & essent. oils.  29-33. 

rasyidah wahab, ahmad asnawi mus, kartini saibeh, shamsul khamis, handry mujih, geofarry gunsalam, dasini, ezron gerald, juhinin sariman, qarin aaron muhammad, qaid ahmad muhammad.  (2021).  soil physico-chemistry in the habitat of rafflesia in kinabalu park, sabah, malaysia..  - journal of tropical biology and conservation.  149-165. 

syarifah haniera sheikh kamal, mohd nazip suratman, shamsul khamis, ahmad najmi nik hassan, mohd syaiful mohammad.  (2021).  host-parasitic relationships between tetrastigma rafflesiae and rafflesia azlanii and rafflesia cantleyi in belum-temenggor forest complex, perak, malaysia.  - pertanika journal of tropical agricultural science.  755-771. 

nor akmalazura jani, farah aina rokman, rashidah iberahim, shamsul khamis.  (2021).  composition and bioactivities of the rhizomes essential oil of curcuma aeruginosa.  - gading journal of science and technology.  66-74. 

muhamad faris osman, soo yee lee, shahrul razid sarbini, siti munirah mohd faudzi, shamsul khamis, badrul hisyam zainudin, khozirah shaari.  (2021).  metabolomics-driven discovery of an introduced species and two malaysian piper betle l. variants.  - plants.  1-19. 

shamsul khamis, nur syamina jamaluddin.  (2021).  an annotated checklist of flowering plants from foothill to seroja camp trail in mount korbu perak semenanjung malaysia.  - seminar saintifik geobiotrail gunung korbu 2021.  183-188. 

s akram, n i ab ghani1,, s khamis and s zulkifly.  (2021).  newly designed chs genic primers for four zingiberaceae species (alpinia mutica, alpinia rafflesiana, hornstedtia leonurus and scaphochlamys kunstleri).  - iop conference series: earth and environmental science.  1-9. 

shamsul, k., nik norafida, n. a., mat salleh, k. and a. latiff.  (2020).  tree species composition, diversity and biomass estimation in lower montane forest at fraser's hill, pahang, peninsular malaysia.  - the malaysian forester.  114-127. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2020).  chemical composition and lipoxygenase activity of the leaves essential oil of rothmannia macrophylla (hook.f.) bremek.  - journal of essential oil bearing plants.  331-336. 

shamsul bin khamis;mohd. nizam bin mohd. said.  (2020).  kajian taksonomi dan ekologi ke atas famili lauraceae di semenanjung malaysia.  - .  . 

shamsul khamis, wan mohd nazri abdul rahman, aiza harun.  (2020).  63 tumbuhan ubatan.  - .  136. 

wan mohd nuzul hakimi wan salleh, shamsul khamis, muhammad nurakmal abdul rahman, mohd azlan nafiah.  (2020).  chemical composition and biological activities of dipterocarpus cornutus dyer essential oil.  - journal of biosciences: zeitschrift für naturforschung c.  171-175. 

a.k. nur hayati, k. shamsul, w.a. wan juliana, m. n. shukor, m. h. shahril, e. alyaa filza.  (2020).  plant community structure and diversity of the rafflesia habitat at the royal belum state park, perak, malaysia.  - the malaysian forester.  387-404. 

muhammad ammar mohd azhar, wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2020).  essential oil composition of three cryptocarya species from malaysia.  - zeitschrift fur naturforschung section c-a journal of biosciences.  297-301. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2020).  chemical composition of the essential oil of diospyros wallichii king & gamble (ebenaceae).  - natural volatiles & essential oils.  12-17. 

norela sulaiman, azman sulaiman, muzzneena ahmad mustapha, izfa riza hazmi, farah shafawati mohd taib, saiful mansor, shamsul khamis, abdullah samat, juliana senawi, salmah yaakop, nurul wahida othman, hani kartini agustar,.  (2020).  majalah knovasi 1/2020.  - pembelajaran aktif di lapangan untuk pemuliharaan kelip-kelip.  1. 

natasa mohd shakri, wan mohd nuzul hakimi wan salleh, shamsul khamis, nor azah mohamad ali, shazlyn milleana shaharudin.  (2020).  chemical composition of the essential oils of four polyalthia species from malaysia.  - zeitschrift fuer naturforschung, section c: a journal of biosciences.  473-478. 

shamsul khamis, kushairi salleh mohd.  (2020).  flora pokok di sekitar pusat penyelidikan bukit fraser, pahang.  - .  19-32. 

natasa mohd shakri, wan mohd nuzul hakimi wan salleh, shamsul khamis, nor azah mohamad ali.  (2020).  chemical characterization of goniothalamus macrophyllus and goniothalamus malayanus leaves essential oils.  - zeitschrift fur naturforschung c a journal of biosciences.  485-488. 

natasa mohd shakri, wan mohd nuzul hakimi wan salleh, shamsul khamis, nor azah mohamad ali, muhammad helmi nadri.  (2020).  composition of the essential oils of three malaysian xylopia species (annonaceae).  - zeitschrift fur naturforschung c a journal of biosciences.  479-484. 

wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2020).  chemical composition and anticholinesterase inhibitory activity of pavetta graciliflora wall. ex ridl. essential.  - zeitschrift fur naturforschung c a journal of biosciences.  467-471. 

norela sulaiman, muzzneena ahmad mustapha, salmah yaakop, azman sulaiman, izfa riza hazmi, nurul wahida othman, juliana senawi, farah shafawati mohd taib, saiful mansor, abdullah samat, hani kartini agustar, shamsul khamis, lailatul nadhirah asri, nur ati.  (2020).  usaha pemuliharaan.  - .  89-119. 

wan mohd nuzul hakimi wan salleh, nur kamilah mohammed, nur athirah hashim, shamsul khamis, salam ahmed abed, saidu jibril.  (2020).  chemical constituents of beilschmiedia penangiana.  - chemistry of natural compounds.  576-577. 

sani aliyu, norazah basar1, yakubu rufai, shamsul khamis.  (2020).  phytochemical analysis and isolation of secondary metabolites from persea declinata.  - malaysian journal of analytical sciences.  970-977. 

syarifah haniera sheikh kamal, mohd nazip suratman, shamsul khamis, ahmad najmi nik hassan, mohd syaiful mohammad.  (2020).  host-parasitic relationships between tetrastigma rafflesiae and rafflesia azlanii and r. cantleyi in belum-temenggor forest complex, perak.  - 7th international conference on science and social research 2020.  1-19. 

wahab, rasyidah, saibeh, k., s. khamis,.  (2020).  tekstur tanah di habitat bunga rafflesia sp. di kawasan taman negara kinabalu, sabah, malaysia.  - the 12th postgraduate symposium,.  20. 

nur syamina jamaluddin, shamsul khamis.  (2020).  a collection of sapotaceae in fraser hill, pahang, peninsular malaysia.  - fraser hill symposium.  1-18. 

shamsul, k., siti nur fateha radzi, siti nur syazwani musthami@sohaimi, nur syamina jamaludin, radhiah zakaria, ahmad fitri, z., mohd nizam, m.s., talip, n., latiff, a..  (2020).  a checklist of tree species in ulu sat forest reserve, kelantan.  - siri kepelbagaian biologi hutan.  132-144. 

k. shamsul, a. norhayati, y.f. ng, a. latiff.  (2020).  watershed.  - .  22-34. 

k. shamsul, a. norhayati, y.f. ng, a. latiff.  (2020).  the flora.  - .  65-96. 

k. shamsul, a. norhayati, y.f. ng, a. latiff.  (2020).  the fauna.  - .  96-121. 

faridah qz, ayuni m. n, shamsul khamis, anisa s. al-hakimi, rosimah nulit, norhidayah mad halid.  (2019).  traditional knowledge on aromatic plants as utilised by the jahai tribe in royal belum state park, perak, malaysia.  - malayan nature journal.  1-8. 

mazlan mohd som, shahril mod husin, aisyah md shukor, alyaa filza effendi, noor ilyani abd rani, shamsul khamis, nur hayati abdul karim, shukor md. nor.  (2019).  rafflesia conservation efforts and management at royal belum state park.  - .  149. 

shamsul k., nizam, m.s., al-hakimi s. anisa, talip n..  (2019).  taxonomic value of petiole anatomy in malaysian litsea complex (lauraceae).  - 11th flora malesiana symposium.  110. 

nur-athirah abdullah, siti nur fatehah radzi, lailatul-nadhirah asri, nor shafikah idris, shahril husin, azman sulaiman, shamsul khamis, norela sulaiman, izfa riza hazmi.  (2019).  insect community in riparian zones of sungai sepetang, sungai rembau and sungai chukai of peninsular malaysia.  - biodiversity data journal.  1-24. 

nur hayati abd karim, shamsul khamis, wan juliana wan ahmad.  (2019).  reassessment of the rafflesia population and microhabitat in royal belum state park, perak.  - international conference on tropical forest science (ictfs): forest for sustainable development goals.  32. 

r siti nur fatehah, k shamsul, w a wan juliana norela sulaiman, m h md. shahril, a d nurul darsani, m s aisah.  (2019).  riverine mangrove plant species composition and vegetation structure of the fireflys habitat at sungai sepetang kampung dew, taiping, perak.  - 14th international annual symposium universiti malaysia terengganu.  47. 

shaharuddin mohamad ismail, shamsul khamis, nurul hidayah shamsul arrifin, faizal mohamad hassan mukhtar.  (2019).  penjelajahan flora nur laman bestari eco resort.  - .  127. 

nur hayati a. k, shamsul k., wan juliana w. a., shahril m. h., alyaa filza e., mohamed shah redza h..  (2019).  re-assessment of the rafflesia populations and habitats in royal belum state park, perak, peninsular malaysia.  - malayan nature journal.  493-498. 

muhamad ikhwanuddin mat esa, rosimah nulit, shamsul khamis, rusea go.  (2019).  ethnomedical survey of plants used by the bateq in kuala koh, kelantan, malaysia.  - malayan nature journal.  439-451. 

shamsul k., m. s. nizam, anisa s. al-hakimi, n. talip.  (2019).  taxonomic value of petiole anatomy in malaysian litsea complex (lauraceae).  - malayan nature journal.  453-465. 

wan mohd nuzul hakimi wan salleh, nur athirah hashim, shamsul khamis.  (2019).  chemical constituents and lipoxygenase inhibitory activity of piper stylosum miq.  - bulletin of the chemical society of ethiopia.  587-592. 

wan mohd nuzul hakimi wan salleh, norkamilah abdullah, nur athirah hashim, heng yen khong, shamsul khamis.  (2019).  aporphine alkaloids from piper erecticaule and acetylcholinesterase inhibitory activity.  - boletin latinoamericano y del caribe de plantas medicinales y aromaticas.  527-532. 

nur athirah hashim, farediah ahmad, wan mohd nuzul hakimi wan salleh, shamsul khamis.  (2019).  a new amide from piper maingayi hk.f. (piperaceae).  - natural product communications.  1-5. 

mohd lip jabit, mohd nazrul hisham daud, shamsul khamis.  (2019).  cytotoxic, nitric oxide and alpha-glucosidase inhibitory activities of biflavonoids from garcinia prainiana king.  - advances in applied chemistry and biochemistry.  55-67. 

ahmad fariz bin mohamed;riza bin sulaiman;shaharudin bin idrus;bisharuzi bin omar;khairul nizam bin abdul maulud;nik mohd noor faizul bin md saad;nor`ain binti mohd jali;rd puteri khairani binti khirotdin;shamsul bin khamis.  (2018).  ukm campus tree id and hazardous trees assessment.  - .  . 

faridah qamaruz zaman, anisa s. al-hakimi, shamsul khamis, fatin f. ruhaizin and syuhada. m. zaidi.  (2018).  anatomy and histochemistry of structures producing aroma in leaves of sygyzium aromaticum (l.) merr. and clausena excavata burm. f..  - pertanika journal of tropical agricultural science.  . 

norela sulaiman, shamsul khamis, azman sulaiman & muzzneena ahmad mustapha.  (2018).  penanaman berembang: usaha pemuliharaan kelip-kelip.  - rampai penyelidikan siri 6. stem untuk komuniti.  . 

nurul ayuni, m.n., faridah, q.z., anisa, s.a., rosimah, n., norhidayah, m.h. & shamsul, k..  (2018).  ethnobotanical documentation of plants used by the jahai tribe in royal belum state park, perak.  - proceedings of the 15th seminar on medicinal and aromatic plants (maps-15).  . 

nik norhazrina nik mohd kamil, ahmad fitri zahari, shamsul khamis, nur aqilah mustafa bakray, haja maideen kader maiden.  (2018).  medicinal plants.  - .  83-90. 

rasyidah wahab, kartini saibeh, shamsul khamis & monica suleiman.  (2018).  kajian komuniti tumbuhan bagi perumah rafflesia (tetrastigma sp.) di lokasi berbeza di taman negara kinabalu, sabah.  - .  . 

soo foong lian, shamsulbin khamis & noraini talip.  (2017).  comparative plant morphology and leaf anatomy of selected accessions of ipomoea batatas (l.) lam. in peninsular malaysia.  - fundamental science congress 2017.  .