siti nurfaeiza abd razak, wan juliana wan ahmad, shamsul khamis, shukor md nor.  (2022).  morpho-physiological strategies of shorea leprosula miq. and shorea acuminata dyer in response to light intensity and nutrient amendments.  - forest.  1-18. 

wan mohd nuzul hakimi wan salleh, natasa mohd shakri, shamsul khamis, william n. setzer, muhammad helmi nadri.  (2022).  chemical composition of three malaysian horsfieldia essential oils.  - natural product research.  1909-1913. 

nor shafikah idris, muzzneena ahmad mustapha, norela sulaiman, shamsul khamis, shahril mohd husin, nurul darsani ahmad darbis.  (2021).  the dynamics of landscape changes surrounding a firefly ecotourism area.  - global ecology and conservation.  1-20. 

muhamad faris osman, soo yee lee, shahrul razid sarbini, siti munirah mohd faudzi, shamsul khamis, badrul hisyam zainudin, khozirah shaari.  (2021).  metabolomics-driven discovery of an introduced species and two malaysian piper betle l. variants.  - plants.  1-19. 

wan mohd nuzul hakimi wan salleh, mohd hafiz arzmi, shamsul khamis, mohd afiq aizat juhari.  (2024).  chemical composition and antioxidant activity of the essential oil of tabernaemontana dichotoma.  - chemistry of natural compounds.  169-170. 

nurunajah ab ghani, wan mohd nuzul hakimi wan salleh, abubakar siddiq salihu, nurulfazlina edayah rasol, shamsul khamis.  (2024).  essential oil composition of antidesma cuspidatum and acetylcholinesterase inhibitory activity.  - chemistry of natural compounds.  167-168. 

roberta chaya tawie tingga, millawati gani, nur azimah osman, nor rahman aifat, eddie chan, shamsul khamis, emelda rosseleena rohani, norlinda mohd-daut, abd rahman mohd ridwan, badrul munir md-zain.  (2024).  interspecific variation in the diet of symphalangus syndactylus and macaca nemestrina at genting highlands, pahang, peninsular malaysia.  - biodiversity data journal.  1-18. 

n.h.a. kadir, w.m.n.h.w. salleh, n.a. ghani, s. khamis, m.a.a juhairi.  (2023).  chemical composition and acetylcholinesterase activity of syzygium pyrifolium (blume) dc. essential oil.  - latin american applied research.  357-359. 

ahmad azrul hakim yahaya, wan mohd nuzul hakimi wan salleh, nurunajah ab ghani, shamsul khamis, nurul syafiqah rezali, mohd afiq aizat juhari.  (2022).  chemical composition and bioactivities of magnolia candollii h.keng essential oil.  - zeitschrift fur naturforschung.  519-523. 

noraini talip, jayasrie eyamalay, mohamad ruzi abdul rahman, mohd nizam mohd said, haja maideen kader maideen, nik norhazrina nik mohd kamil, nur aqilah mustafa bakray, shamsul khamis, ahmad fitri zohari, nur farah ain zainee, hamidun bunawan.  (2024).  ciri adaptasi anatomi daun bougainvillea spectabilis w. yang dijangkiti virus bougainvillea chlorotic vein-banding.  - malaysian applied biology.  43-54. 

wan mohd nuzul hakimi wan salleh, mohd hafiz arzmi, shamsul khamis, mohd afiq aizat juhari.  (2024).  chemical composition and antioxidant activity of the essential oil of tabernaemontana dichotoma.  - chemistry of natural compounds.  169-170. 

roberta chaya tawie tingga, millawati gani, nur azimah osman, nor rahman aifat, eddie chan, shamsul khamis, emelda rosseleena rohani, norlinda mohd-daut, abd rahman mohd ridwan, badrul munir md-zain.  (2024).  interspecific variation in the diet of symphalangus syndactylus and macaca nemestrina at genting highlands, pahang, peninsular malaysia.  - biodiversity data journal.  1-18. 

nurunajah ab ghani, wan mohd nuzul hakimi wan salleh, abubakar siddiq salihu, nurulfazlina edayah rasol, shamsul khamis.  (2024).  essential oil composition of antidesma cuspidatum and acetylcholinesterase inhibitory activity.  - chemistry of natural compounds.  167-168. 

seemab akram, shahrizim zulkifly, shamsul khamis, nurul izza ab ghani.  (2023).  preliminary in silico analysis of chs1 gene in commelinids clade: family zingiberaceae, costaceae, and poaceae.  - pertanika j. trop. agric. sci..  425-437. 

s akram, n i ab ghani1,, s khamis and s zulkifly.  (2021).  newly designed chs genic primers for four zingiberaceae species (alpinia mutica, alpinia rafflesiana, hornstedtia leonurus and scaphochlamys kunstleri).  - iop conference series: earth and environmental science.  1-9. 

siti munirah mat yunoh, nur hayati abd. karim, halimah alias, mohd. noor mat isa, wan kiew lian, shamsul khamis, mohd afiq aizat juhari.  (2023).  rafflesia di pahang.  - .  21. 

wee suk ling, siti munirah mat yunoh, nur hayati abd. karim, shamsul khamis.  (2023).  rafflesia di pahang.  - .  22. 

mohd. hizamri mohd. yasin, norela sulaiman, shamsul khamis.  (2023).  rafflesia di pahang; rafflesia of pahang.  - .  1-69. 

norela sulaiman, muzzneena ahmad mustapha, salmah yaakop, azman sulaiman, izfa riza hazmi, nurul wahida othman, juliana senawi, farah shafawati mohd taib, saiful mansor, abdullah samat, hani kartini agustar, shamsul khamis, lailatul nadhirah asri, nur ati.  (2020).  kelip-kelip anugerah alam di waktu malam.  - .  30. 

shamsul khamis, wan mohd nazri abdul rahman, aiza harun.  (2020).  63 tumbuhan ubatan.  - .  136. 

siti munirah mat yunoh, nur hayati abd. karim, halimah alias, mohd. noor mat isa, wan kiew lian, shamsul khamis, mohd afiq aizat juhari.  (2023).  rafflesia di pahang.  - .  21. 

wee suk ling, siti munirah mat yunoh, nur hayati abd. karim, shamsul khamis.  (2023).  rafflesia di pahang.  - .  22. 

shamsul, k., siti nur fateha radzi, siti nur syazwani musthami@sohaimi, nur syamina jamaludin, radhiah zakaria, ahmad fitri, z., mohd nizam, m.s., talip, n., latiff, a..  (2020).  hutan ulu sat: nadi pemeliharaan alam semulajadi kelantan (ulu sat forest: the heart of kelantan's nature conservation).  - .  13. 

shamsul khamis, kushairi salleh mohd.  (2020).  bukit fraser kepelbagaian biology dan persekitaran fizikal.  - .  13. 

k. shamsul, a. norhayati, y.f. ng, a. latiff.  (2020).  biodiversity of water catchment forest: hutan simpan bukit cherakah, selangor.  - .  12. 

mohd. hizamri mohd. yasin, norela sulaiman, shamsul khamis.  (2023).  rafflesia di pahang; rafflesia of pahang.  - .  1-69. 

shamsul khamis, noralasikin yahya, rukayah aman.  (2022).  kamus herba fitoterapi.  - kamus fitoterapi.  1-61. 

shamsul khamis, wan mohd nazri abdul rahman, aiza harun.  (2020).  63 tumbuhan ubatan.  - .  136. 

shaharuddin mohamad ismail, shamsul khamis, nurul hidayah shamsul arrifin, faizal mohamad hassan mukhtar.  (2019).  penjelajahan flora nur laman bestari eco resort.  - .  127. 

mazlan mohd som, shahril mod husin, aisyah md shukor, alyaa filza effendi, noor ilyani abd rani, shamsul khamis, nur hayati abdul karim, shukor md. nor.  (2019).  rafflesia conservation efforts and management at royal belum state park.  - .  149. 

shamsul khamis mohd. afiq aizat juhari, siti nurfaeiza abd razak.  (2023).  hubungkait perumah rafflesia (tetrastigma sp.) dengan komuniti tumbuhan dan faktor tanah di hutan simpan jerantut tambahan, mukim ulu cheka, daerah jerantut, pahang.  - seminar ekspedisi saintifik kepelbagaian biologi hutan ulu cheka, jerantut, pahang..  . 

norela bt. sulaiman;siti norhafizah binti ahmad tarmidzi;mohd afiq aizat bin juhari;shamsul bin khamis.  (2022).  highland stingless bee identification and the assessment of domesticated stingless bees at high altitude.  - .  . 

muzzneena binti ahmad mustapha;norela bt. sulaiman;shamsul bin khamis.  (2022).  mapping potential firefly habitats using texture based classification of satellite imagery.  - .  . 

nur syamina jamaluddin, shamsul khamis.  (2020).  a collection of sapotaceae in fraser hill, pahang, peninsular malaysia.  - fraser hill symposium.  1-18. 

shamsul bin khamis;mohd. nizam bin mohd. said.  (2020).  kajian taksonomi dan ekologi ke atas famili lauraceae di semenanjung malaysia.  - .  .